Keeping in mind I'm not a tech, so only basic explainations available.
So it looks like no current through Q4. Am I right in thinking that with a PNP, if b is -ve relative to e, there should be current?
R12 Vdrop confirms no current flow through Q4.
You are correct regarding PNP transistors. The arrow indicates the direction of the voltage drop, assuming normal forward bias operation.
Typically, the voltage drop is about 0.6-0.7V. Your measurements show 1.3V. As you increase the Vbe, the transistor works harder/higher
current. Maybe the higher Vbe caused Q4 to "burn out"??? Doubtful, Q4e voltage is higher than expected because of no current flow and
no associated Vdrop across R12, which will probably be 400mV like R9 giving a reasonable(?) Q4 Vbe.
I can't reconcille the no current through R12 other than Q4 suspect faulty. With current flow through Q4 it will hopefully correct Q5 Vbe
turning it on so there is current flow down R17,18,19,,, branch.
Mouser has MPSA13 on back order, expected delivery about Sept 13. If you need something sooner I could check my spares, worst case
remove a Q4 from one of my units. I'm away from home maybe 10 days so no real advantage I think, maybe best to wait, install Q4 and
problem remains.... Maybe someone more knowledgable will chime in and confirm we are on the right/wrong track.