Well the topology for sure. Haven't dug into it. But The 555a uses a real old-school single rail voltage system as does the 505, 777. It is also capacitor coupled and uses all NPN output trannies - a so called quasi complementary output. I would imagine the 5500 is dual rail (diff. amp), direct coupled, and full complementary - in short a modern amp ;-)
The old cap coupled amps do have a special sound to them. I don't know if it is better per say, but very musical and open. I like them a lot.